Six mutants, including bsx14, which had very low fertility (Table 1), did not transmit the trait, at least within the limits of the tested progeny populations. Alterations of GSF can be generated by pollen irradiation and result in the restoration of the ancestral hermaphroditic trait. 1991) and achieved by alterations of reproductive organ formation at early stages of development (Grantet al. Thus, the arrest of carpel development in the male flower is the earliest event in the establishment of the sexual dimorphism in white campion (Figure 1). The identified modifiers can act semidominantly and are under developmental control. alba) is a biennial or short lived perennial in the pink (Caryophyllaceae) family.